Tennessee’s woodland vole records number 823, split between a small photographed set and a much larger body of specimen and survey data. That puts the species in the occasional bucket for Tennessee mammals, meaning it turns up more predictably than most small mammals but still isn’t among the state’s most frequently recorded rodents.
From limestone glades to mountain duff
The state’s clearest clusters sit in two very different landscapes. Around Nashville, Radnor Lake State Natural Area and Cedars of Lebanon State Park both host the vole in the thin, rocky soil of the Central Basin’s eastern redcedar glades, ground that looks nothing like typical woodland. Farther east, the species turns up again in the deep leaf litter and loamy soil of the Great Smoky Mountains’ forest floor. What ties the two together isn’t the surface vegetation but what’s underneath it. Any soil loose enough to tunnel through and hold a network of shallow runways will do, whether that’s glade dirt over limestone or mountain forest duff.
A vole that rarely surfaces
Unlike most voles, this species spends nearly all its time below ground. It digs shallow tunnels through loose topsoil and leaf litter, gnawing on roots, bark, and bulbs, and pushing up small mounds of loosened dirt the way a mole does. That underground habit keeps it out of sight even in places where it’s recorded often, and it’s why sightings tend to cluster around burrow openings and pushed-up soil rather than the animal itself.
Status in Tennessee
The species is native to Tennessee. NatureServe ranks it S5, secure, the highest rank on the scale. Tennessee’s records peak each spring, heaviest in April, then fall off through summer before another rise in fall, a pattern that likely tracks breeding activity and surface digging as much as the animal’s actual numbers.



