California vole

Microtus californicus

Also known as: Amargosa Vole

LCLeast Concern

California voles are compact grassland rodents that build runways under thick stems instead of crossing open ground in long leaps. Their small ears, short tail, and blunt face suit a life spent close to cover. The species belongs mainly to California's coast, valleys, and nearby uplands, where periodic surges in numbers can strongly affect predators, plants, farms, and disease cycles.

Range through the year

Where California vole is seen

California vole is recorded in 2 US states from 9,533 occurrence records, most heavily in California (9,461 records) and Oregon (72).

Current range9,533 records · 2 states
AlabamaArizonaArkansasCalifornia · 9,461 records · view state pageColoradoConnecticutDelawareDistrict of ColumbiaFloridaGeorgiaIdahoIllinoisIndianaIowaKansasKentuckyLouisianaMaineMarylandMassachusettsMichiganMinnesotaMississippiMissouriMontanaNebraskaNevadaNew HampshireNew JerseyNew MexicoNew YorkNorth CarolinaNorth DakotaOhioOklahomaOregon · 72 records · view state pagePennsylvaniaRhode IslandSouth CarolinaSouth DakotaTennesseeTexasUtahVermontVirginiaWashingtonWest VirginiaWisconsinWyomingAlaskaAKHawaiiHIPuerto RicoPR
Occurrence data by state and month
California vole – occurrence records by US state and month (9,533 total across 2 states)
StateJanFebMarAprMayJunJulAugSepOctNovDecYear
California8594789701,2311,1181,0669226245905814575659,461
Oregon919722813·1·1172

Based on 12,064 georeferenced US records, 2000-2026. Shading is observation density (where people record sightings), not true abundance. Alaska, Hawaii & Puerto Rico shown as insets.

Explore by state

California vole across 2 states

Occurrence records, conservation status, and the places it turns up, state by state. Open a state for detail, or visit its full page.

  • CaliforniaNativeCommon11,990 records

    State rankUnranked

    Notable placesGolden Gate National Recreation Area · Point Reyes National Seashore · Bodega Marine Reserve · Russell Research Station (UC)

    Countiesrecorded in 12

    Year-round in California, with recorded sightings peaking in March–July, with a smaller rise in January.

  • OregonNativeOccasional73 records

    State rankApparently Secure

    Notable placesSiskiyou National Forest · Cascade-Siskiyou National Monument · Denman Wildlife Area

    Countiesrecorded in 6

    Year-round in Oregon, with recorded sightings peaking in June.

IUCN Red List status

  1. LC
  2. NT
  3. VU
  4. EN
  5. CR
  6. EW
  7. EX

LC – Least Concern

Widespread and abundant; not at risk.

Elevation

From 200 georeferenced records, the California vole is recorded from sea levelsea level to 2,342 m7,684 ft, with half of sightings below 36 m118 ft.

Each band shows the share of georeferenced sightings that fall within it. Records cluster where people live and hike, so low, populated elevations are over-represented. Read this as observation density, not true abundance.

Best encounter

Where and when to see a California vole

Your best chance: California.

Where
CaliforniaCA
Trophic relationshipsFood web What a California vole eats, and what eats it.
Eaten by6
Northern Rubber BoaCharina bottaeGiant Garter SnakeThamnophis gigasRed Diamond RattlesnakeCrotalus ruberCalifornia Ground SquirrelOtospermophilus beecheyiAmerican Barn OwlTyto furcataLong-tailed WeaselNeogale frenata
This speciesCalifornia vole
Clustered GoldenweedPyrrocoma racemosaYerba MansaAnemopsis californicaCoast Live OakQuercus agrifoliaOregon OakQuercus garryanaCalifornia Black OakQuercus kelloggii
Eats5

A sample of documented feeding relationships, filtered to species that also occur in the US. Diet and predators vary by region and season, and this is not a complete list, many animals eat, and are eaten by, more than what has been formally recorded here.

Life history

The California vole (43–47 g1.5–1.7 oz) is a herbivore that first breeds around 0.1 years, carries a 21-day gestation, and bears about 4.6 young per litter. Maximum recorded longevity is 2.1 years.

Life cycle

  1. Gestation21 days
  2. Birthlitter of 4.6 young
  3. Sexual maturity0.1 yr (female) · 0.1 yr (male)
  4. Typical lifespan2.1 years
  5. Maximum recorded longevity2.1 years

Reproduction

Gestation
21days
Litter size
4.6youngaverage
Litters / year
4.92litters
Age at first breeding
0.1years♀ 0.1 · ♂ 0.1 yr

Diet composition

Herbivore

By foraging composition: 80% plant material, 20% seeds & grain.

  • Plant material80%
  • Seeds & grain20%

Most active around the clock.

Typical values for the species, not local measurements. Maximum longevity is the longest lifespan on record; most individuals live far less.

Commonly confused with

Animals people sometimes mistake for the california vole, and how to tell them apart.

Life inside a grass tunnel

A California vole looks rounder and shorter-tailed than a deer mouse. Its ears barely rise above the fur. It clips paths through grass and uses nests hidden at ground level or just below it. Those covered lanes let the vole feed while staying less visible to owls, hawks, foxes, snakes, and many other hunters.

Moist cover in a dry region

Meadows, marsh edges, weedy fields, and grassy openings provide dense food and concealment. Most records fall within California. Reports from outside the accepted range need careful review because related voles look similar. Clusters near cities and universities also reflect where people search, so they aren’t a direct measure of population density.

Fast breeding and uneven years

Voles eat grasses, sedges, roots, and other green plants. Females can breed quickly when rain and fresh growth create good conditions. Numbers may climb and fall sharply between years. That pulse feeds predators and changes grazing pressure. It also explains why a familiar field may seem full of runways one season and quiet later.

Small animal, broad effects

California voles can damage crops and young plantings, yet they are a major link between vegetation and predators. Burrowing mixes shallow soil, and clipped plants alter the ground layer. The species isn’t considered threatened overall. Wetland drainage, mowing, fire, and development can still remove individual colonies or shift them into narrow margins.

Data & sources

The data and visualizations on this page draw on these open sources: